Joma, A., and W. P. Mackay. “Revision of the African Ants of the Bothroponera Pumicosa Species Complex (Hymenoptera: Formicidae: Ponerinae)”. Sociobiology, vol. 62, no. 4, Dec. 2015, pp. 538-63, doi:10.13102/sociobiology.v62i4.845.